Anadyricarpa altingiosimila N. Maslova et Herman, Ettingshausenia louravetlanica (Herman et Shczepetov) Herman et Moiseeva  


Anadyricarpa circled center right.



Grebenka, Yelisseev Locality, Site 11c

Locality Map

Geological Map

Sedimentary Log



Description of Anadyricarpa altingiosimila N. Maslova et Herman taken from Maslova and Herman (2004).

The infructescence includes a relatively massive axis, with longitudinal ribs and a maximal diameter equal to 2.5 mm and several alternatively attached capitate infructescences with a diameter of about 8 mm. The maximal observed number of heads per axis is nine. Closer to the axis base, the heads are attached with a pedicel (up to 4 mm long). Toward the apex of the axis, the pedicels become shorter, and apical heads are sessile.
The head consists of a central core (3 mm in diameter); radially attached, tightly adpressed flowers of various maturity; and fruits. Rounded or oval scars of slightly different diameter are scars of shed fruits and carpels: larger scars are obviously remains of mature fruits. The mean number of flowers per head is fewer than 35.

The microstructure of particular flowers was studied with SEM. The length of a separate flower is 1900- 2100 µm; the width varies from 230 to 750 µm depending on the maturity of the carpel. Each individual flower consists of a well-developed perianth and a solitary carpel. The members of the perianth form a tube, which embraces the carpel along nearly its entire length. In underdeveloped narrow carpels only apices of short stylodes remain free. The perianth tube persists in mature fruits as well. The number of perianth members and degree of perianth differentiation were not revealed because of the insufficient preservation. The upper surface of the flower is folded. In transverse section, a relatively thick layer is visible that consists of perianth members, which envelop the gynoecium along its entire length. The upper cuticle of the perianth is formed by elongate cells with greater cutinized longitudinal walls.

The mature fruit is narrowly elliptical, about 1700 µm long and up to 700 µm wide, with a distinct longitudinal suture. The fruit base is conical. All visible fruits are devoid of stylodes. However, it is still uncertain whether this is taphonomic or whether the stylodes were shed after fruit became mature. The cuticle of the fruit wall is formed by tetragonal cells, often with equal sides.

The following description of Ettingshausenia louravetlanica is taken from Moiseeva (2010).

The leaves are simple, entire margined, with small lateral lobes or without lobes, medium and large in size. The leaf blade is often asymmetrical, of variable outlines: oval, oval–ovate, broadly rhomboidal, or obovate. The petiole is long, more than 7 cm in length. The leaf base is broadly or narrowly cuneate and decurrent. The apices of the central and lateral lobes are acute or obtuse. The lateral lobes are lacking or small, triangular, often are situated at different height, and differ in size. The leaf margin is dentate-emarginate, with small glands terminating the teeth. The venation is palmate-pinnate and craspedodromous. The basal veins often come from the midrib asymmetrically; they are alternating or, more rarely, opposite, situated at a distance of 5–25 mm from the leaf base. The basal veins are nearly as thick as other secondary veins and have one or two acroscopic and five to seven basiscopic deviations. There are up to four infrabasal veins, developed in various degree. The tertiary venation varies from orthogonal-reticulate to branchy scalariform.

The species resembles the type species E. cuneifolia (Bronn) Stiehle from the Cenomanian of Czech Republic (Peruc-Korycany Formation) by the rhomboidal leaf blade, small lateral lobes, and finely dentate leaf margin (Kvacek and Váchová, 2006). E. louravetlanica differs from this species, as well as from the majority of other species of the genus, by the asymmetrical leaf base and leaf blade: arrangement and size of lateral lobes and asymmetrical (alternating) deviation of basal veins. Moreover, the above described leaves of E. louravetlanica are distinctive by their tertiary venation, which is a combination of orthogonal-reticulate and branchy-scalariform types, contrasting to the majority of other species characterized by branchy-scalariform tertiary venation.


The heads of Anadyricarpa altingiosimila were found in association with the leaves of Ettingshausenia louravetlanica and bark fragments. Bark fragments vary in outline and size. Their longitudinal relief is virtually identical to that of extant Platanus.

This photograph was made in the field. The specimen is in the collection of the Geological Institute, Russian Academy of Sciences, Moscow, Russia. For further details of the context of the collection see Spicer et al. (2002).